Vorlage:Elder et al., 1993
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- Elder, J.F. Turner, B. Thomerson, J.E. & Taphorn, D.C. 1993. Karyotypes of nine Venezuelan annual killifishes (Cyprinodontidae), with comments on karyotype differentiation in annual killifishes. Ichthyological Exploration of Freshwaters, 4 (3): 261-268.