Marcelo R. de Carvalho

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Marcelo Rodrigues de Carvalho

  • Departamento de Zoologia, Instituto de Biociências, Universidade de São Paulo, Rua do Matão, Trav. 14, no. 101, São Paulo, SP, CEP 05508-090, Brazil.
  • E-mail:

Beschriebene Taxa



  • Carvalho, M.R.d. 1999. A systematic revision of the electric ray genus Narcine Henle, 1834 (Chondrichthyes: Torpediniformes: Narcinidae), and the higher-level phylogenetic relationships of the orders of elasmobranch fishes (Chondrichthyes). Unpublished PhD thesis, City University of New York, 735 pp. Zitatseite 


  • Carvalho, M.R. 2001. A new species of electric ray, Narcine leoparda, from the tropical eastern Pacific Ocean (Chondrichthyes: Torpediniformes: Narcinidae). Proceedings of the Biological Society of Washington, 114 (3): 561-573. (BHL) Zitatseite 


  • McEachran, J.D. & Carvalho, M.R. 2002. Gymnuridae: Butterfly Rays. Pp. 575–577 in:
  • Carpenter, K.E. (Ed.) 2002. The Living Marine Resources of the Western Central Atlantic. Volume 1: Introduction, Mollusks, Crustaceans, Hagfishes, Sharks, Batoid fishes and Chimaeras. FAO, Rome. Zitatseite  Zitatseite 
  • Carvalho, M.R., Stehmann, M.F.W. & Manilo, L.G. 2002. Torpedo adenensis, a New Species of Electric Ray from the Gulf of Aden, with Comments on Nominal Species of Torpedo from the Western Indian Ocean, Arabian Sea, and Adjacent Areas (Chondrichthyes: Torpediniformes: Torpedinidae). American Museum Novitates, 3369: 1-34. (PDF) Zitatseite 



  • Carvalho, M.R., Maisey, J.G. & Grande, L. 2004. Freshwater stingrays of the Green River Formation of Wyoming (Early Eocene), with the description of a new genus and species and an analysis of its phylogenetic relationships (Chondrichthyes: Myliobatiformes). Bulletin of the American Museum of Natural History, 284: 1–136. (doi) Zitatseite 



  • Carvalho, M.R.d. 2010. Morphology and phylogenetic relationships of the giant electric ray from the Eocene of Monte Bolca, Italy (Chondrichthyes: Torpediniformes). Pp. 183-198 in: Elliott, D.K., Maisey, J.G., Yu, X. & Miao, D. (eds): Morphology, Phylogeny and Paleobiogeography of Fossil Fishes. Verlag Dr. Friedrich Pfeil, München. Zitatseite 


  • Carvalho, M.R. & Lovejoy, N.R. 2011. Morphology and phylogenetic relationships of a remarkable new genus and two new species of Neotropical freshwater stingrays from the Amazon basin (Chondrichthyes: Potamotrygonidae). Zootaxa, 2776: 13-48. (doi) Zitatseite 
  • Carvalho, M.R., Sabaj Pérez, M.H. & Lovejoy, N.R. 2011. Potamotrygon tigrina, a new species of freshwater stingray from the upper Amazon basin, closely related to Potamotrygon schroederi Fernandez-Yépez, 1958 (Chondrichthyes: Potamotrygonidae). Zootaxa, 2827: 1-30. Zitatseite 


  • Ebert, D.A., White, W.T., Ho, H.-C., Last, P.R., Nakaya, K., Séret, B., Straube, N., Naylor, G.J.P. & Carvalho, M.R. 2013. An annotated checklist of the chondrichthyans of Taiwan. Zootaxa, 3752 (1): 279–386. (doi) Zitatseite 
  • Loboda, T.S. & Carvalho, M.R. 2013. Systematic revision of the Potamotrygon motoro (Müller & Henle, 1841) species complex in the Paraná-Paraguay basin, with description of two new ocellated species (Chondrichthyes: Myliobatiformes: Potamotrygonidae). Neotropical Ichthyology, 11 (4): 693-737. (doi) Zitatseite 
  • Vaz, D.F.B. & Carvalho, M.R. 2013. Morphological and taxonomic revision of species of Squatina from the Southwestern Atlantic Ocean (Chondrichthyes: Squatiniformes: Squatinidae). Zootaxa 3695: 1–81. Zitatseite 


  • Fontenelle, J.P., Silva, J.P.C.B. & Carvalho, M.R. 2014. Potamotrygon limai sp. nov., a new species of freshwater stingray from the upper Madeira River system, Amazon basin (Chondrichthyes: Potamotrygonidae). Zootaxa, 3765 (3): 249-268. (doi) Zitatseite 
  • Rosa, R.S., Gama, C., Fontenelle, J.P., Carvalho, M.R. de, Lasso, C.A. & Morales-Betancourt, M.A. 2014. Potamotrygon scobina. Pp. 256–260 in: Lasso, C.A., Rosa, R.S., Sanchez-Duarte, P., Morales-Betancourt, M.A. & Agudelo-Córdoba, E. (Eds.): Rayas de Agua Dulce (Potamotrygonidae) de Suramérica. Parte I. Colombia, Venezuela, Ecuador, Perú, Brasil, Guyana, Surinam y Guayana Francesa: Diversidad, Bioecología, Uso y Conservación. Serie Recursos Hidrobiológicos y Pesqueros Continentales de Colombia. IX. Instituto de Investigación de los Recursos Biológicos Alexander von Humboldt, Bogotá. Zitatseite 


  • Ebert, D.A., Haas, D.L. & Carvalho, M.R. de 2015: Tetronarce cowleyi, sp. nov., a new species of electric ray from southern Africa (Chondrichthyes: Torpediniformes: Torpedinidae). Zootaxa,, 3936 (2): 237-250. Zitatseite 
  • Fontenelle, J.P. & Carvalho, M.R. 2015. Systematic implications of brain morphology in Potamotrygonidae (Chondrichthyes: Myliobatiformes). Journal of Morphology, 277 (2): 252–263. (doi) Zitatseite 
  • Jones, C.M., Driggers, W.B. III, Castro, J.I. & Carvalho, M.R. de 2015. On the attribution of authorship for several elasmobranch species in Müller and Henle’s Systematische Beschreibung der Plagiostomen (Chondrichthyes, Elasmobranchii). Zootaxa, 4052 (5): 569-572. Zitatseite 
  • Silva, J.P.C.B. & Carvalho, M.R. 2015: Systematics and morphology of Potamotrygon orbignyi (Castelnau, 1855) and allied forms (Chondrichthyes: Myliobatiformes: Potamotrygonidae). Zootaxa, 3982 (1): 1-82. (Abstract). Zitatseite 


  • Carvalho, M.R. de 2016. Potamotrygon rex, a new species of Neotropical freshwater stingray (Chondrichthyes: Potamotrygonidae) from the middle and upper rio Tocantins, Brazil, closely allied to Potamotrygon henlei (Castelnau, 1855). Zootaxa, 4150 (5): 537-565. Zitatseite 
  • Carvalho, M.R. de. 2016. Description of two extraordinary new species of freshwater stingrays of the genus Potamotrygon endemic to the rio Tapajós basin, Brazil (Chondrichthyes: Potamotrygonidae), with notes on other Tapajós stingrays. Zootaxa, 4167 (1): 1-63. (PDF) Zitatseite 
  • Carvalho, M.R. 2016. Neotropical stingrays: Family Potamotrygonidae. Pp.619-655 in:
  • Last, P.R., White, W.T., Carvalho, M.R., Séret, B., Stehmann, M.F.W. & Naylor, G.J.P. (eds.) 2016. Rays of the World. CSIRO Publishing, Comstock Publishing Associates. i-ix + 1-790. Zitatseite  Zitatseite 
  • Carvalho, M.R.d., Rosa, R.S. & Araújo, M.L.G. 2016. A new species of Neotropical freshwater stingray (Chondrichthyes: Potamotrygonidae) from the Rio Negro, Amazonas, Brazil: the smallest species of Potamotrygon. Zootaxa, 4107 (4): 566-586. Zitatseite 
  • Carvalho, M.R.d., Loboda, T.S. & Silva, J.P.C.B.d. 2016. A new subfamily, Styracurinae, and new genus, Styracura, for Himantura schmardae (Werner, 1904) and Himantura pacifica (Beebe & Tee-Van, 1941) (Chondrichthyes: Myliobatiformes). Zootaxa, 4075 (3): 201-221. doi: 10.11646/zootaxa.4175.3.1 Zitatseite 
  • Carvalho, M.R., Last, P.R. & Séret, B. 2016. 18. Torpedo Rays. Family Torpedinidae. Pp. 184-203 in:
  • Last, P.R., White, W.T., Carvalho, M.R., Séret, B., Stehmann, M.F.W. & Naylor, G.J.P. (eds.) 2016. Rays of the World. CSIRO Publishing, Comstock Publishing Associates. i-ix + 1-790. Zitatseite  Zitatseite 
  • Carvalho, M.R. de & White, W.T. 2016. Narcine baliensis, a new species of electric ray from southeast Asia (Chondrichthyes: Torpediniformes). Zootaxa, 4127 (1): 149-160. Zitatseite 
  • Last, P.R., White, W.T., Carvalho, M.R., Séret, B., Stehmann, M.F.W. & Naylor, G.J.P. (eds.) 2016. Rays of the World. CSIRO Publishing, Comstock Publishing Associates. i-ix + 1-790. Zitatseite 
  • Viana, S.T.d.F. & Carvalho, M.R.d. 2016. Redescription of Squalus acutipinnis Regan, 1908, a Valid Species of Spiny Dogfish from Southern Africa (Chondrichthyes: Squaliformes: Squalidae). Copeia, 104 (2): 539–553. (Abstract) doi: 10.1643/CI-14-217 Zitatseite 
  • Viana, S.T.d.F., Carvalho, M.R.d. & Gomes, U.L. 2016. Taxonomy and morphology of species of the genus Squalus Linnaeus, 1758 from the Southwestern Atlantic Ocean (Chondrichthyes: Squaliformes: Squalidae). Zootaxa, 4133 (1): 1-89. Zitatseite 


  • Fontenelle, J.P. & Carvalho, M.R.d. 2017. Systematic revision of the Potamotrygon scobina Garman, 1913 species-complex (Chondrichthyes: Myliobatiformes: Potamotrygonidae), with the description of three new freshwater stingray species from Brazil and comments on their distribution and biogeography. Zootaxa, 4310 (1): 1-63. (doi) Zitatseite 
  • Viana, S.T.d.F.L., Lisher, M.W. & Carvalho, M.R.d. 2017. Two new species of short-snouted dogfish sharks of the genus Squalus Linnaeus, 1758, from southern Africa (Chondrichthyes: Squaliformes: Squalidae). Marine Biodiversity, [1-28] First Online: 08 April 2017. (doi) Zitatseite 
  • Viana, S.T.d.F.L., Carvalho, M.R.d. & Ebert, D.A. 2017. Squalus bassi sp. nov., a new long-snouted spurdog (Chondrichthyes: Squaliformes: Squalidae) from the Agulhas Bank. Journal of Fish Biology, 91 (4): 1178–1207. (doi) Zitatseite 
  • Yokota, L. & Carvalho, M.R.d. 2017. Taxonomic and morphological revision of butterfly rays of the Gymnura micrura (Bloch & Schneider 1801) species complex, with the description of two new species (Myliobatiformes: Gymnuridae). Zootaxa, 4332 (1): 1-74. (doi) Zitatseite 


  • de Figueiredo Petean, F. & Carvalho, M.R. 2018. Comparative morphology and systematics of the cookiecutter sharks genus Isistius Gill (1864) (Chrondrichthyes: Squaliformes: Dalatiidae). PLoS ONE, 13 (8): e0201913. (doi) Zitatseite 
  • Vaz, D.F.B. & Carvalho, M.R. 2018. New Species of Squatina (Squatiniformes: Squatinidae) from Brazil, with Comments on the Taxonomy of Angel Sharks from the Central and Northwestern Atlantic. Copeia, 106 (1): 144-160. (doi) Zitatseite 



  • Soares, K.D.A. & Carvalho, M.R. 2020. Phylogenetic relationship of catshark species of the genus Scyliorhinus (Chondrichthyes, Carcharhiniformes, Scyliorhinidae) based on comparative morphology. Zoosystematics and Evolution, 96 (2): 345-395. (doi) Zitatseite 
  • Viana, S.T.F.L. & Carvalho, M.R. 2020. Squalus shiraii sp. nov. (Squaliformes, Squalidae), a new species of dogfish shark from Japan with regional nominal species revisited. Zoosystematics and Evolution, 96 (2): 275-311. (doi) Zitatseite 


  • Loboda, T.S., Lasso, C.A., Rosa, R.S. & Carvalho, M.R. 2021.: Two new species of freshwater stingrays of the genus Paratrygon (Chondrichthyes: Potamotrygonidae) from the Orinoco basin, with comments on the taxonomy of Paratrygon aiereba. Neotropical Ichthyology, 19 (2): e200083. (doi) Zitatseite 


  • Carvalho, M.R. 2022. Family Torpedinidae, Torpedo rays (Pp. 543-547, Pls. 52-54); family Narcinidae, Numbfishes (Pp. 547-552, Pls. 54-55); family Narkidae, Sleeper rays (Pp. 552-557, Pls. 55-56). In:
  • Heemstra, P.C., Heemstra, E., Ebert, D.A., Holleman, W. & Randall, J.E. (eds.) 2022. Coastal fishes of the western Indian Ocean. Volume 1. South African Institute for Aquatic Biodiversity, Makhanda, South Africa: i-ii, 1-637, i-v, Pls. 1-82. Zitatseite  Zitatseite