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  • Weiss, S., Knizhin, I., Kirillov, A. & Froufe, E. 2006. Phenotypic and genetic differentiation of two major phylogeographical lineages of arctic grayling Thymallus arcticus in the Lena River, and surrounding Arctic drainages. Biological Journal of the Linnean Society, 88 (4): 511-525. (doi)