Vorlage:Spies et al., 2011

• Spies, I.B., Stevenson, D.E., Orr, J.W. & Hoff, G.R. 2011. Molecular systematics of the skate subgenus Arctoraja (Bathyraja: Rajidae) and support for an undescribed species, the leopard skate, with comments on the phylogenetics of Bathyraja. Ichthyological Research, 58 (1): 77-83. (doi)